1. Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China
2. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China
3. Sino-Danish College of University of Chinese Academy of Sciences, Beijing 101400, China
4. Sino-Danish Center for Education and Research, Beijing 101400, China
5. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, China