1. Geomicrobiology, Center for Applied Geosciences, University of Tuebingen, 72076 Tuebingen, Germany
2. Delaware Environmental Institute, University of Delaware, Newark, Delaware 19716, United States
3. Department of Environmental Earth System Science, Stanford University, Stanford, California 94305, United States
4. School of Earth Sciences, University of Bristol, Queens Road, Bristol BS8 1QU, United Kingdom