1. Section for Microbial Ecology and Biotechnology, Department of Plant and Environmental Sciences, University of Copenhagen, Thorvaldsenvej 40, DK-1871 Frederiksberg C, Denmark
2. Key Laboratory of Urban Environment and Health, Institute of Urban Environment, Chinese Academy of Science, 1799 Jimei Road, Xiamen 361021, China
3. Institute of Environmental Microbiology, College of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou 350002, Fujian China
4. State Key Laboratory of Urban and Regional Ecology, Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China