1. Department of Civil and Environmental Engineering, KAIST, Daejeon 34141, South Korea
2. Microorganism Resources Division, National Institute of Biological Resources, NIBR, Incheon 22689, South Korea
3. Center for Environmental Biotechnology, Department of Microbiology, Department of Civil and Environmental Engineering, Department of Biosystems Engineering and Soil Science, University of Tennessee, Knoxville, Tennessee 37996, United States
4. Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee 37831, United States