1. Department of Biological Science, College of Science, Sungkyunkwan University, Suwon 16419, South Korea
2. Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystems/College of the Environment and Ecology, Xiamen University, Xiamen 361102, China
3. Institute of Integrated Science and Technology, Nagasaki University, Nagasaki 852-8521, Japan
4. Organization for Marine Science and Technology, Nagasaki University, Nagasaki 852-8521, Japan
5. Department of Marine Science, College of Nature Science, Incheon National University, Incheon 22012, South Korea