1. School of Marine Sciences, Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, Pearl River Estuary Marine Ecosystem Research Station, Ministry of Education, Sun Yat-sen University; Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China
2. School of Ecology, Sun Yat-sen University, Guangzhou 510275, China