1. Division
of Life Science, The Hong Kong University of Science and Technology (HKUST), Clear Water Bay, Kowloon, Hong Kong
2. HKUST Shenzhen Research Institute, Shenzhen 518057, China
3. Key Laboratory of the Coastal & Wetland Ecosystems, Ministry of Education, College of the Environment & Ecology, Xiamen University, Xiamen, Fujian 361102, China