1. Institute for Marine and Antarctic Studies, 20 Castray Esplanade, Battery Point, Tasmania 7004, Australia
2. Institute for Marine and Antarctic Studies, University of Tasmania, School Road, Newnham, Tasmania 7248, Australia
3. Central Science Laboratory, University of Tasmania, Private Bag 74, Hobart, Tasmania 7001, Australia
4. Tasmanian School of Medicine, University of Tasmania, Hobart, Tasmania 7000, Australia