1. BioTechnology Institute, University of Minnesota, St. Paul, Minnesota 55108, United States
2. Department of Microbiology & Immunology, University of Minnesota, Minneapolis, Minnesota 55455, United States
3. Department of Surgery, University of Minnesota, Minneapolis, Minnesota 55455, United States
4. Department of Soil, Water & Climate, and Department of Microbial and Plant Biology, University of Minnesota, St. Paul, Minnesota 55108, United States