1. School of Biosciences, Cardiff University, Museum Avenue, Cardiff CF10 3AX, U.K.
2. Centre for Environment, Fisheries and Aquaculture Science (Cefas), Suffolk, Lowestoft NR33 0HT, U.K.
3. Environment Agency, Red Kite House, Howbery Park, Wallingford, Oxfordshire OX10 8BD, U.K.
4. Lancaster Environment Centre, UK Centre for Ecology and Hydrology, Library Avenue, Bailrigg, Lancaster LA1 4AP, U.K.