1. School of Marine Sciences, Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, Pearl River Estuary Marine Ecosystem Research Station, Ministry of Education, Sun Yat-Sen University, Southern Marine Science and Engineering Guangdong Laboratory, Zhuhai 519082, China
2. Department of Biology, Center for Environment, Biodiversity and Conservation, The University of Texas at Tyler, Tyler, Texas 75799, United States
3. Key Laboratory of Tropical Marine Bio-Resources and Ecology, Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
4. School of Ecology, Sun Yat-Sen University, Guangzhou 510275, China