1. College of Life Sciences, Fujian Normal University, Fuzhou, Fujian 350117, China
2. College of Food and Bioengineering, Fujian Polytechnic Normal University, Fuqing 350300, China
3. State Key Laboratory of Microbial Technology, School of Life Science, Shandong University, Jinan 250100, China
4. Department of Chemistry and Biochemistry, Ohio University, Athens, Ohio 45701-2978, United States
5. Engineering Research Center of Industrial Microbiology of Ministry of Education, Fujian Normal University, Fuzhou, Fujian 350117, China