1. Department of Natural Resource Sciences, McGill University, Ste. Anne de Bellevue, Quebec H9X 3V9, Canada
2. School of Environmental Studies, University of Victoria, Victoria, British Columbia V8P 5C2, Canada
3. Procellaria Research & Consulting, Victoria, British Columbia V9A 5C3, Canada
4. Environment and Climate Change Canada, Pacific Wildlife Research Centre, Delta, British Columbia V4K 3N2, Canada
5. Department of Biology, University of Western Ontario, London, Ontario N6A5B7, Canada
6. Environment and Climate Change Canada, Saskatoon, Saskatchewan S7N 3H5, Canada
7. National Wildlife Research Centre, Environment and Climate Change Canada, 1125 Colonel By Dr, Ottawa, Ontario K1S 5B6, Canada