1. Graduate School of Global Environmental Studies, Kyoto University, Kyoto 606-8501, Japan
2. Regional Environment Conservation Division, National Institute for Environmental Studies, Ibaraki 305-8506, Japan
3. Graduate School of Human and Environmental Studies, Kyoto University, Kyoto 606-8316, Japan
4. Faculty of Environmental Science and Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Nagasaki 852-8521, Japan