1. State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou 570228, China
2. Center for Eco-Environment Restoration of Hainan Province & Key Laboratory of Agro-Forestry Environmental Processes and Ecological Regulation of Hainan Province, School of Environment and Ecology, Hainan University, Haikou 570228, China
3. Key Laboratory of Microbial Technology for Industrial Pollution Control of Zhejiang Province, College of Environment, Zhejiang University of Technology, Hangzhou 310032, China
4. Hainan Academy of Ocean and Fisheries Sciences, Haikou 571126, China
5. Haereticus Environmental Laboratory, P.O. Box 92, Clifford, Virginia 24533, United States
6. Lee Kong Chian Natural History Museum, National University of Singapore, Singapore 117377, Singapore
7. Tropical Marine Science Institute, National University of Singapore, Singapore 119227, Singapore
8. Department of Biological Sciences, National University of Singapore, Singapore 117558, Singapore