1. Fujian Provincial Key Laboratory for Coastal Ecology and Environmental Studies, State Key Laboratory of Marine Environmental Science, College of the Environment and Ecology, Xiamen University, Xiamen 361102, Fujian, China
2. Grantham Institute – Climate Change and the Environment and Department of Life Sciences, Imperial College London, Exhibition Road, London SW7 2AZ, U.K.
3. GEOMAR Helmholtz Centre for Ocean Research Kiel, Kiel D-24148, Germany