1. Department of Ocean Science, Division of Life Science and Hong Kong Branch of the Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), The Hong Kong University of Science and Technology, Kowloon, Hong Kong 999077, China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, Guangdong 510642, China
3. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9, Canada
4. Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Alberta T6G 2E3, Canada