1. Key Laboratory of Pollution Ecology and Environmental Engineering, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang, Liaoning 110016, China
2. University of Chinese Academy of Sciences, Beijing 100039, China
3. Department of Microbiology, Department of Civil and Environmental Engineering, The University of Tennessee, Knoxville, Tennessee 37996, United States
4. Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee 37831, United States
5. Department of Biosystems Engineering and Soil Science, Center for Environmental Biotechnology, The University of Tennessee, Knoxville, Tennessee 37996, United States