1. Université de Lyon, Université Claude Bernard Lyon 1, CNRS, ENTPE, UMR 5023 LEHNA, F 69622 Villeurbanne, France
2. School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, U.K.
3. Centre for Environment Fisheries and Aquaculture Science, Pakefield Road, Lowestoft, Suffolk NR33 0HT, U.K.