1. School of Earth, Ocean and Environment, University of South Carolina, Columbia, South Carolina 29208-0001, United States
2. Belle W. Baruch Institute for Marine and Coastal Sciences, University of South Carolina, South Carolina 29208, United States
3. North Inlet-Winyah Bay National Estuarine Research Reserve, Georgetown, South Carolina 29440, United States
4. Marine, Earth and Atmospheric Sciences, North Carolina State University, Raleigh, North Carolina 27695, United States