1. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China
2. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China
3. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, Iowa 50011, United States
4. The James Hutton Institute, Dundee, Scotland DD2 5DA, U.K.