1. Department of Biology and Evolution of Marine Organisms, Stazione Zoologica Anton Dohrn, Villa Comunale, Naples 80121, Italy
2. Department of Physical, Earth and Environmental Sciences, University of Siena, Via Mattioli, 4, Siena 53100, Italy
3. Department of Integrative Marine Ecology, Stazione Zoologica Anton Dohrn, Villa Comunale, Naples 80121, Italy