1. School of Biological and Environmental Sciences, Liverpool John Moores University, 3 Byrom Sreett, Liverpool L3 3AF, U.K.
2. Queen’s University Marine Laboratory, Queen’s University Belfast, 12-13 The Strand, Portaferry, BT22 1PF, Northern Ireland U.K.
3. Department of Animal Ecology, Federal Institute of Hydrology, Am Mainzer Tor 1, 56068 Koblenz, Germany
4. British Antarctic Survey, High Cross Madingley Road, Cambridge, CB3 0ET, U.K.