1. Shandong Key Laboratory of Water Pollution Control and Resource Reuse, School of Environmental Science and Engineering, Shandong University, Qingdao, Shandong 266237, P. R. China
2. The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, College of Life Science, Shandong University, Qingdao, Shandong 266237, P. R. China
3. College of Life and Environmental Sciences, University of Exeter, Exeter EX4 4QD, Devon, U.K.