1. State Key Laboratory of Soil and Sustainable Agriculture, Institute of Soil Science, Chinese Academy of Sciences, 71 East Beijing Road, Nanjing 210008, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. School of Biological Sciences, University of Aberdeen, Aberdeen AB24 3UU, U.K.
4. Key Laboratory of Watershed Geographic Sciences, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, China