1. Center for Research on Sustainable Forests, University of Maine, Orono, Maine 04469, United States
2. School of Forest Resources, University of Maine, Orono, Maine 04469, United States
3. School of Marine Sciences, University of Maine, Orono, Maine 04469, United States
4. School of Environmental and Forest Sciences, University of Washington, Seattle, Washington 98195, United States
5. Department of Earth and Environmental Science, University of Exeter, Penryn Campus, Cornwall TR10 9FE, U.K.