1. Depatment of Biological and Agricultural Engineering, North Carolina State University, Raleigh, North Carolina 27695, United States
2. Department of Biology and Institute for Great Lakes Research, Central Michigan University, Mount Pleasant, Michigan 48859, United States
3. Pasture Systems and Watershed Management Research Unit, Agricultural Research Service, United States Department of Agriculture, University Park, Pennsylvania 16802, United States
4. Department of Civil and Environmental Engineering, Pennsylvania State University, University Park, Pennsylvania 16802, United States
5. Department of Biological and Environmental Engineering, Cornell University, Ithaca, New York 14853, United States