1. Odum School of Ecology and the Savannah River Ecology Laboratory, University of Georgia, 140 E. Green Street, Athens, Georgia 30602-0002, United States
2. Institute for Environmental and Spatial Analysis, Watkins Academic Building, University of North Georgia, 3820 Mundy Mill Road, Gainesville, Georgia 30503-1358, United States
3. Crop and Soil Sciences, University of Georgia, Carlton Street, Athens, Georgia 30602-0002, United States
4. Odum School of Ecology, University of Georgia, 140 E. Green Street, Athens, Georgia 30602-0002, United States