1. State Key Laboratory for Biology of Plant Diseases and Insect Pests, Institute of Plant Protection, Chinese Academy of Agricultural Sciences, Beijing 100193, P. R. China
2. Department of Plant, Soil and Microbial Sciences, Michigan State University, East Lansing, Michigan 48824, United States
3. Department of Environment and Geography, University of York, Heslington, York, YO10 5NG, United Kingdom