1. College of Resources and Environment, Huazhong Agricultural University, Wuhan430070, China
2. State Key Laboratory of Agricultural Microbiology, Huazhong Agricultural University, Wuhan430070, China
3. School of Fisheries, Aquaculture and Aquatic Sciences, Auburn University, Auburn, Alabama36849, United States
4. Agriculture Microbiology Department, Faculty of Agriculture, Benha University, Moshtohor, Qalubia13736, Egypt