1. Marine Laboratory, Duke University, Beaufort, North Carolina 28516, United States
2. Program in Environmental Health and Toxicology, Duke University, Durham, North Carolina 27708, United States
3. Key Laboratory of Marine Environment and Ecology (Ministry of Education), College of Environmental Science and Engineering, Ocean University of China, Qingdao, 266003, P. R. China
4. Biology Department, Duke University, Durham, North Carolina 27708, United States
5. Civil and Environmental Engineering, Duke University, Durham, North Carolina 27708, United States