1. Department of Biological Science, University of Lethbridge, Lethbridge, Alberta T1K 3M4, Canada
2. Department of Environmental Sciences, College of the Coast and Environmental, Louisiana State University, Baton Rouge, Louisiana 70803, United States
3. Institut des Sciences de la Mer de Rimouski, Université du Québec à Rimouski, Rimouski, Québec G5L 3A1, Canada