1. School of Environmental Sciences, University of Liverpool, Liverpool L69 3GP, U.K.
2. UK Centre for Ecology & Hydrology, Bush Estate, Penicuik EH26 0QB, U.K.
3. Atmosphere and Ocean Research Institute, University of Tokyo, Kashiwa, Chiba 277-8564, Japan
4. UK Centre for Ecology & Hydrology, Lancaster Environment Centre, Library Avenue, Bailrigg, Lancaster LA1 4AP, U.K.