1. Department of Physical and Environmental Sciences, University of Toronto Scarborough, 1265 Military Trail, Scarborough, Ontario M1C 1A4, Canada
2. School of Mathematics and Statistics, Rochester Institute of Technology, 85 Lomb Memorial Dr., Rochester, New York 14623, United States
3. Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks St, Toronto, Ontario M5S 3B2, Canada