1. Advanced Water Management Centre, The University of Queensland, St. Lucia, Brisbane, Queensland 4072, Australia
2. School of Environmental Science and Engineering, Shandong University, Qingdao, Shandong 266237, China
3. Australian Centre for Ecogenomics, The University of Queensland, St. Lucia, Brisbane, Queensland 4072, Australia
4. School of Municipal and Environmental Engineering, Harbin Institute of Technology, Harbin 150090, China
5. Microbiology, IWWR, Faculty of Science, Radboud University Nijmegen, Heyendaalseweg 135, 6525 AJ Nijmegen, The Netherlands