1. Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai 264003, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Department of Bioengineering, School of Marine Science and Technology, Harbin Institute of Technology, Weihai 264209, China
4. Laboratory of Microbial Ecology and Matter Cycles, School of Marine Sciences, Sun Yat-Sen University, Zhuhai 519082, China
5. Key Laboratory of Marine Chemistry Theory and Technology, Ministry of Education, Ocean University of China, Qingdao 266100, China