1. Department of Ecosystem Science and Management, University of Wyoming, Laramie, Wyoming 82071, United States
2. Department of Life & Environmental Sciences and Sierra Nevada Research Institute, University of California, Merced, California 95343, United States
3. Smithsonian Tropical Research Institute, Apartado 0843-03092 Balboa, Ancon, Panama
4. Department of Geography, University of California, Santa Barbara, California 93106, United States
5. Canadian Light Source, University of Saskatchewan, Saskatoon, Canada S7N 0X4