1. State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou, Hainan 570228, P. R. China
2. Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, P. R. China
3. Department of Environmental Science & Engineering, Fudan University, Shanghai 200438, P. R. China
4. Department of Biology/Aquatic Ecology, Ecology Building, Lund University, Lund SE-22100, Sweden