1. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China
2. Department of Ocean Science, The Hong Kong University of Science and Technology, Clear Water Bay, Kowloon, Hong Kong, China
3. SZU-HKUST Joint PhD Program in Marine Environmental Science, Shenzhen University, Shenzhen 518060, China
4. Department of Chemical and Biological Engineering, The Hong Kong University of Science and Technology, Kowloon, Hong Kong, China
5. Institute for Advanced Study, Shenzhen University, Shenzhen 518060, China