1. National Agricultural Research Center for Kyushu Okinawa Region, National Agriculture and Food Research Organization, 496 Izumi, Chikugo, Fukuoka 833-0041, Japan
2. National Food Research Institute, National Agriculture and Food Research Organization, 2-1-18 Kannondai, Tsukuba, Ibaraki 305-8642, Japan
3. National Institute of Crop Science, National Agriculture and Food Research Organization, 2-1-12 Kannondai, Tsukuba, Ibaraki 305-8642, Japan