1. Department of Bioscience, Faculty of Bioresources, Mie University, 1515 Kamihama, Tsu, Mie 514-8507, Japan; National Institute of Agrobiological Resources, 2-1-2 Kannondai, Tsukuba, Ibaraki 305-0856, Japan; and Division of Biofunctional Materials, Department of Fundamental Science, Research Institute for Advanced Science and Technology, Osaka Prefecture University, 1-2 Gakuen-cho, Sakai, Osaka 599-8570, Japan