1. Institute for Human Science & Biomedical Engineering, National Institute of Advanced Industrial Science and Technology, Central 6, 1-1-1 Higashi, Tsukuba, Ibaraki 305-8566, Japan, and National Institute of Advanced Industrial Science and Technology, Laboratory of Gene Function Analysis, Institute of Molecular and Cell Biology, Central 2, 1-1-1 Umezono, Tsukuba, Ibaraki 305-8566, Japan