1. Biotechnology Research Center, The University of Tokyo, Tokyo 113-8657, Japan
2. Faculty of Pharmaceutical Sciences, Hokkaido University, Sapporo 060-0812, Japan
3. Collaborative Research Institute for Innovative Microbiology, The University of Tokyo, Tokyo 113-8657, Japan
4. National Institute of Advanced Industrial Science and Technology (AIST), Tokyo 135-0064, Japan