1. Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin, 300308, China
2. Key Laboratory of Systems Microbial Biotechnology, Chinese Academy of Sciences, Tianjin, 300308, China
3. University of Chinese Academy of Sciences, Beijing, 100049, China
4. Department Applied and Molecular Microbiology, Institute of Biotechnology, Technische Universität Berlin, Berlin, 13355, Germany