1. Department of Systems Biology, Columbia University, New York, New York 10027, United States
2. Integrated Program in Cellular, Molecular, and Biomedical Studies, Columbia University, New York, New York 10027, United States
3. School of Physics and Astronomy, University of Minnesota, Minneapolis, Minnesota 55455, United States
4. Department of Pathology and Cell Biology, Columbia University, New York, New York 10027, United States