1. Division of Quantitative Biology, National Institute for Basic Biology, National Institutes of Natural Sciences, Okazaki 444-8787, Japan
2. Quantitative Biology Research Group, Exploratory Research Center on Life and Living Systems (ExCELLS), National Institutes of Natural Sciences, Okazaki 444-8787, Japan
3. Graduate School of Science, Nagoya City University, Nagoya 467-8501, Japan
4. Department of Basic Biology, Faculty of Life Science, SOKENDAI (Graduate University for Advanced Studies), Okazaki 444-8787, Japan