1. Department of Molecular Biotechnology, Graduate School of Advanced Sciences of Matter, Hiroshima University, Higashi-Hiroshima, Hiroshima 739-8530, Japan
2. Advanced Low Carbon Technology Research and Development Program, Japan Science and Technology Agency (JST-ALCA), Chiyoda-ku, Tokyo 102-0076, Japan
3. Department of Bioscience, Tokyo University of Agriculture, Tokyo 156-8502, Japan