1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao 266237, PR China
2. National Glycoengineering Research Center, Shandong University, Qingdao 266237, PR China
3. CAS Key Lab of Biobased Materials, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao 266101, PR China