1. Biosciences, Geoffrey Pope Building, College of Life and Environmental Sciences, University of Exeter, Exeter EX4 4QD, U.K.
2. Biodomain, Shell Technology Center Houston, 3333 Highway 6 South, Houston, Texas 77082-3101, United States
3. School of Natural and Environmental Sciences, Devonshire Building, Faculty of Science, Agriculture and Engineering, Newcastle University, Newcastle-upon-Tyne NE1 7RU, U.K.