1. Graduate Program in Biophysical Sciences, University of Chicago, Chicago, Illinois 60637, United States
2. Department of Chemistry, University of Chicago, Chicago, Illinois 60637, United States
3. Center for Physics of Evolving Systems and Department of Biochemistry and Molecular Biology, University of Chicago, Chicago, Illinois 60637, United States
4. Pritzker School of Molecular Engineering, University of Chicago, Chicago, Illinois 60637, United States